Trackhouse upstaged the factory Aprilias to take its first-ever MotoGP race 1-2 in the Assen sprint, with Raul Fernandez leading home Ai Ogura.

Neither works Aprilia even made the podium, as polesitter Jorge Martin slipped right back to fifth behind team-mate Marco Bezzecchi.

Despite Aprilia's practice domination and top-four grid sweep, the RS-GPs did not have the race pace to escape.

For most of the distance, all four Aprilias, the Ducatis of VR46' Fabio Di Giannantonio and works pair Marc Marquez and Pecco Bagnaia, plus Tech3 KTM's Enea Bastianini all ran together in an eighth-bike lead train.

Di Giannantonio was the main early mover in this and looked like he may even come through to give Ducati an unlikely win at one stage.

He and the Trackhouse pair were able to break away from the rest when Martin and Bezzecchi lost several seconds in a big intra-team fight over fourth that Bezzecchi won. Both lost ground and positions during the race with crucial mistakes.

Though Di Giannantonio got onto leader Fernandez's tail, Ogura had superior pace in the closing laps and was able to demote the Ducati and ensure the Trackhouse 1-2.

Bezzecchi's fourth place inched his championship lead back up to nine points over Martin, who fended off big Bagnaia attacks for fifth.

Bagnaia had come back through after a bad start and some very tight racing with Marquez - which turned out to be in vain as a post-race penalty for going over track limits in his last lap fight with Martin dropped Bagnaia back behind Marquez to seventh.

Bastianini completed the scorers in eighth.

Pedro Acosta would have been among this group had he not run wide mid-race. He recovered to ninth.

Fabio Quartararo held on after his strong qualifying to complete the top 10 on his Yamaha.

With Joan Mir crashing out on lap one, LCR's Diogo Moreira was the best Honda in 11th.